Crown stratification ratio models for Tectona grandis L. f in Oluwa Forest reserve, Nigeria


  • Omobolanle Temitope Faleye
  • Lucas Aderemi Akomolede
  • Olalekan Kehinde Ajayi
  • Opeyemi Philips Akinsulire



crown ratio, modeling, Oluwa forest reserve, Tectona grandis, tree growth


This research investigated crown ratio models for Tectona grandis plantation in Oluwa Forest reserve (Ondo State, Nigeria) using variables of slenderness coefficient and merchantable height. With three non-linear regression functions - logistic, Chapman-Richard and exponential we showed that basal area, tree stem volume and mean tree height per hectare were of high significance. In the same vein, there were fewer tree species in the class of higher diameter and height than there were in lower class. There were also more trees in the co-dominant and intermediate classes than in the dominant and suppressed layers. The lack of emergence in the plantation reflected the past disturbance of the ecosystem. Most of the tree growth variables were significantly different in different canopy layers in the study area. Based on the evaluation models, the three functions investigated for tree crown ratio modeling gave
constant and reliable results in all canopy layers considering their indices. Especially, Chapman-Richard and exponential functions gave consistent trends and good fits for crown ratio models. It is recommended to put strict measures in place to avert any level of illegalities that may likely disrupt the delicate equilibrium of the ecosystem. It is also recommended that complexity revealed in this study is sustained in the region, and encouraged in other parts of Nigeria.


Aber, J.D., 1979. Foliage-height profiles and succession in northern hardwood forests. Ecology, 60, 18-23. DOI:

Adejuwon, J.O., 1971. Agricultural colonization in twentieth century in Western Nigeria: Distribution and vegetation characteristics. Journal of Tropical Geography, 39, 1-10.

Adekunle, V.A.J., Akindele, S.O., Fuwape, J.A., 2004. Structures and yield models for tropical lowland rainforest ecosystem of south west Nigeria. Food, Agriculture and Environment, 2, 395-399.

Adekunle, V.A.J., Ige, P.O., 2006. Logging and logging residues of some selected economic tropical hardwood timber species in free area forest of Ondo State, Nigeria. Journal of Applied Tropical Agriculture, 11 (2), 88-99.

Adekunle, V.A.J., 2007. Non-linear regression models for timber volume estimation in natural forest ecosystem, southwest Nigeria. Research Journal of Forestry, 1 (2), 40-50. DOI:

Adekunle, V.A.J., Olagoke, A.O., 2008. Diversity and bio-volume of tree species in natural forest ecosystem in the bitumen-producing area of Ondo State, Nigeria: a baseline study. Biodiversity and Conservation, 17, 2735-2755. DOI:

Adesoye, P.O., Oluwadare, A.O., 2008. Interim crown ratio models for a mixed Tectona grandis and Gmelina arborea stand in the University of Ibadan, Nigeria. Research Journal of Forestry, 2 (1), 34-42. DOI:

Akinyemi, O.D., Ugbogu, O.A., Adedokun, D., Sefiu, H., Odewo, T.K., Odofin, B.T., Ibidapo, V., 2002. Floristic study of Onigambari lowland rainforest reserve. In: Abu, J.E., Oni, P.I., Popoola, L. (eds) Forestry and challenges of sustainable livelihood. Proceeding of the 28th Annual Conference of

the Forestry Association of Nigeria, Akure, 4th-8th November, 2002, 346-357.

Alder, D., Abayomi, J.O., 1994. Assessment of data requirements for sustained yield calculations. Unpublished report prepared for the Nigeria Tropical Action Program, FORMECU, Federal Department of Forestry, Ibadan, Nigeria, 28p.

Avery, T., Burkhart, H.E., 2001. Forest Measurements. 5th ed. McGraw- hill, New York. Bond, B.J., Franklin, J.F., 2002. Aging in Pacific Northwest forests: a selection of recent research. Tree Physiology, 22, 73-76. DOI:

Brown, M.J., Parker, G.G., 1994. Canopy light transmittance in a chronosequence of mixed-species deciduous forests. Canadian Journal of Forest Research, 24, 1691-1703. DOI:

Chazdon, R.L., Brancalion, P.H.S., Laestadius, L., Bennett-Curry, A., Buckingham, K., Kumar, C., Moll-Rocek, J., Vieira I.C.G., Wilson, S.J., 2016. When is a forest a forest? Forest concepts and definitions in the era of forest and landscape restoration. Ambio, 45 (5), 538-550. DOI:

Chukwu, O., Dau, J.H., Ogundipe, O.C., Wali, M.D., 2018. Crown ratio assessment for Proposis africana (Guill. and Perr.) Taub species in Makurdi, Nigeria. Journal of Research in Forestry, Wildlife & Environment, 10 (1): 20-27.

Clutter, J.L., Fotson, J.C., Pienaar, L.V., Brister, G.H., Bailey, R.L., 1983. Timber Management: A Quantitative Approach. John Wiley and sons, New York.

Cole, W., Lorimer, C.G., 1994. Predicting tree growth from crown variables in managed Northern hardwood stands, Forest Ecology and Management, 67, 159-175. DOI:

Connell, J.H., Slatyer, R.O., 1977. Mechanisms of succession in natural communities and their role in community stability and organization. The American Naturalist, 111 (982), 1119-1144. DOI:

Dyer, M., Burkhart, H., 1987. Compatible crown ratio and crown height models, Canadian Journal of Forest Research, 17, 572-574. DOI:

Fiala, A.C.S., 2003. Forest canopy structure in western Oregon: Characterization, methods for estimation, prediction, and importance to avian species. Corvallis, OR: Oregon State University. 335p. M.S. Thesis.

Franklin, J.F., Spies, T.A., Van Pelt, R., Carey, A.B., Thornburg, D.A., Berg, D.R., Lindenmayer, D.B., Harmon, M.E., Keeton, W.S., Shaw, D.C., Bible, K., Chen, J., 2002. Disturbances and structural development of natural forest ecosystems with silvicultural implications, using Douglas-fir forests as an example. Forest Ecology and Management, 155, 399-423. DOI:

Hardiman, B.S., Bohrer, G., Gough, C.M., Vogel, C.S., Curtisi, P.S., 2011. The role of canopy structural complexity in wood net primary production of a maturing northern deciduous forest. Ecology, 92 (9), 18-27. DOI:

Hann, D.W., 1997. Equation for predicting the largest crown width of stand growth trees in Western Oregon. Forestry research laboratory, research contribution 17. Oregon State University, Corvallis. Hanus, M.I., Hann, D.W., Marshal, D.D., 2000. Predicting Height to Crown Base for Undamaged and Damaged Trees, Southwestern Oregon, research contribution 29. Oregon State University, Corvallis. Hasenauer, H., Monserud R., 1996. A crown ratio model for Austrian forests. Forest Ecology and Management, 84, 49-60.

Henderson, J.A., 1981. Succession of two habitat types in western Washington. In: Means, J., ed. Forest succession and stand development research in the Northwest. Proceedings of the Symposium. Oregon State University, Corvallis, 80-86.

Holdaway, M., 1986. Modeling tree crown ratio. The Forestry Chronicle, 451-455. DOI:

Iloeje, N.P., 1981. A new geography of Nigeria (A new revised edition, published in Great Britain by B. Williams). Colwes Ltd, London, pp. 85-120.

Kershaw, J.A., Maguire, D.A., Hann, D.W., 1990. Longevity and Duration of Radial Growth in Douglasfir Branches. Canadian Journal of Forest Research, 20, 1690-1695. DOI:

Kozlowski, T., Kramer, P., Pallardy. S., 1991. The Physiological Ecology of Woody Plants. Academic Press, New York. Lowman, M.D., Moffett, M.W., 1993. The ecology of tropical rain forest canopies. Trees, 8, 104-107. DOI:

Marshall, D.D., Johnson, G.P., Hann, D.W., 2003. Crown Profile Equations for Stand Grown Western Hemlock in Northwestern Oregon. Canadian Journal of Forest Research, 33, 2059-2066. DOI:

Mcgraughey, R.J., 1997. Visualizing forest stand dynamics using the stand visualization system. In: Proceedings of the 1997 ACSM/ASPRS Annual Convention and Exposition, April 7-10, 1997.

Seatle, WA. American Society for Photogrammetry and Remote Sensing, 4, 248-257.

Monserud, R.A., Sterba, H., 1996. A basal area increment model for individual trees growing in even - and uneven aged forest stands in Austria. Forest Ecology and Management, 80, 57-80. DOI:

Naesset, E., Okland, T., 2002. Estimating tree height and tree crown properties using airborne scanning laser in a boreal nature reserve. Remote Sensing of Environment, 79, 105-115. DOI:

Navaratil, S., 1997. Wind damage thinned stands. In: Proceedings of a commercial thinning workshop. Whitecourt, AB, October 17-18, 1997, pp. 29-36.

Nigerian Meteorological Society (NIMS), 2007. Proceedings of the international conference on the impact of extreme weather in Africa. 11-15 November, 2007, Akure, Nigeria. Ojo, L.O., 2004. The fate of the tropical rainforest in Nigeria. Abeku Sector of Omo Forest Reserve.

Global Nest: The International Journal, 6 (2), 116-130.

Oliver, C.D., 1981. Forest development in North America following major disturbances. Forest Ecology and Management, 3, 153-168. DOI:

Oliver, C.D., Larson, B.D., 1990. Forest stands dynamics. McGraw-Hill, New York. Osunade, M.A.A., 1991. Agricultural change by supplanting process in a traditional farming system. International Journal of Ecology and Environmental Sciences, 17, 201-209.

Parker, G.G., 1995. Structure and microclimate of forest canopies. Pages 73-106. In: M. Lowman and N. Nadkarni, eds. Forest canopies: a review of research on a biological frontier. Academic Press, San Diego,

Popoola, F.S., Adesoye, P.O., 2012. Crown ratio models for Tectona grandis (Linn. f) stands in Osho forest reserve, Oyo State, Nigeria. Journal of Forest and Environmental Science, 28 (2), 63-67. DOI:

Reineke, L.H., 1933. Perfecting a stand-density index for even-aged forest. Journal of Agricultural Research, 46, 627-638.

Schomaker, M.E., Zarnoch, S.J., Bechtold, W.A., Latelle, D.J., Burkman, W.G., Cox, S.M., 2007. Crown-condition classification: A guide to data collection and analysis Gen. Tech. Rep. SRS-102. DOI:

Asheville, N.C: U.S. Department of Agriculture, Forest Service, Southern Research Station. 78p. Smith III, T.J., 1988. Differential distribution between subspecies of the mangrove Ceriops tagal: Competitive interactions along a salinity gradient. Aquatic Botany, 32(1-2), 79-89. DOI:

Soares, P., 1999. Modelação Do Crescimento Da Árvore Em Eucaliptais Em 1ª Rotação Localizados Nas Regiões Norte E Centro Litoral, PhD Thesis, ISA, Lisboa, 1999, 369p.

Soares, P., Tome, M., 2001. A tree crown ratio prediction equation for eucalyptus plantations. Annals of Forest Science, 58 (2), 193-202. DOI:

Stewart, G.H., 1988. The influence of canopy cover on understorey development in forests of the western Cascade Range, Oregon, USA. Vegetation, 76, 79-88. DOI:

Temesgen, H., Lemay, V., Mitchell, S.J., 2005. Tree crown ratio models for multi-species and multilayered stands of Southeastern British Columbia. The Forest Chronicle, 81, 1. DOI:

Van Pelt, R., Franklin, J.F., 2000. Influence of canopy structure on the understory environment in tall, old-growth, conifer forests. Canadian Journal of Forest Research, 30, 1231-1245. DOI:

Vanclay, J.K., 1994. Modeling forest growth and yield. Application of mixed tropical forests. CAB International, Wallingford U.K. 329p

Waide, R.B., 2008. Tropical rainforest. In: Encyclopedia of ecology (2nd edition), 2, 679-683. DOI:

Waring, R.H., Schlesinger, W.H., 1985. Forest ecosystems: concepts and management. Academic Press, Orlando, Fla. Zamora, B.A., 1981. Understory development in forest succession: an example from the inland Northwest. In: Means, J., ed. Forest succession and stand development research in the northwest. Proceedings of the Symposium. Corvallis, OR: Oregon State University, 63-79.






Original Research Paper

How to Cite

Temitope Faleye, O., Aderemi Akomolede, L., Kehinde Ajayi, O., & Philips Akinsulire, O. (2021). Crown stratification ratio models for Tectona grandis L. f in Oluwa Forest reserve, Nigeria. Acta Biologica Slovenica, 64(1), 70-90.

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